EDUCATION

MD, University of Turku, Finland  1981;
PhD, University of Turku 1985;

Docentships:
•Medical Biochemistry, University of Turku 1990
•Anatomy, University of Oulu, Finland 1993
•Cell and developmental biology, University of Oulu 2009

Postdoc & foreign visits

Jefferson Institute of Molecular Medicine, Department of Dermatology, Thomas Jefferson University, Philadelphia, PA, USA, 1987-1990